Reference

Glossary of Terms

A comprehensive terminology reference for lichen identification: from growth forms and reproductive structures to chemical tests and anatomical features.

Lichens have a rich and specialized vocabulary. Whether you are working through a dichotomous key, reading a species description, or studying lichen biology, understanding these terms is essential. This glossary covers the terminology used across lichen identification, anatomy, ecology, and chemistry, organized with visual references for key structures followed by a complete alphabetical listing.

Visual Guide to Key Structures

The most important morphological terms illustrated with photographs. Click through to the full definitions in the alphabetical glossary below.

Crustose lichen

Crustose

Flat crust fused to the surface; cannot be removed without damaging substrate

Foliose lichen

Foliose

Leafy growth form with distinct upper and lower surfaces

Fruticose lichen

Fruticose

Shrubby or hair-like three-dimensional growth form

Squamulose lichen

Squamulose

Small, overlapping, scale-like lobes

Gelatinous lichen

Gelatinous

Jelly-like form that swells dramatically when wet

Apothecia on lichen

Apothecia

Open disc-shaped sexual fruiting bodies that produce spores

Soredia on lichen

Soredia

Powdery asexual propagules lacking a cortex

Isidia on lichen

Isidia

Corticated finger-like outgrowths for asexual reproduction

Podetia on Cladonia

Podetia

Stalked vertical structures characteristic of Cladonia

Lirellae on lichen

Lirellae

Elongated line-like fruiting bodies

Rhizines on lichen underside

Rhizines

Root-like attachment structures on foliose lichen undersides

Pseudocyphellae on lichen

Pseudocyphellae

Pores in the cortex appearing as white spots or lines

Cyphellae on lichen

Cyphellae

Cup-shaped pores on the lower surface

Umbilicate lichen

Umbilicate

Attached by a single central point, like rock tripe

Alphabetical Glossary

A comprehensive reference for the terminology used in lichen identification, biology, and ecology. Terms are organized alphabetically.

Apothecia
ApotheciaOpen, cup- or disc-shaped sexual fruiting bodies that produce spores. Singular: apothecium.
Areoles
Small, defined patches on a crustose lichen surface, separated by cracks.
Ascospore
Spore produced sexually inside an ascus.
Ascus
Sac-like structure inside apothecia where spores form. Plural: asci.
Basidiomycete yeasts
Yeasts discovered in 2016 living in the cortex of many macrolichens; potential third symbiotic partner.
Byssoid
Cotton-like or wispy growth form.
Cephalodia
Specialized structures containing cyanobacteria in lichens that also have green algae as primary photobiont (e.g., Lobaria pulmonaria).
Cilia
Hair-like projections from the margin of lichen lobes.
Coralloid
Branching like coral; describes a type of isidium.
Cortex
Dense, protective outer layer of fungal tissue on the lichen surface.
Corticolous
Growing on bark.
Crustose
CrustoseGrowth form adhering tightly to substrate like a crust; cannot be removed without damaging the substrate.
Cyphellae
CyphellaeSmall, cup-shaped pores on the underside, found in Sticta.
Dichotomous
Branching into two equal parts; a common pattern in fruticose lichens.
Doctrine of Signatures
Historical belief that organisms resemble the body part they can heal.
Epiphytic
Growing on the surface of plants, but not parasitic.
Esorediate
Lacking soredia.
Filamentous
Thread-like growth form.
Foliose
FolioseLeafy growth form with distinct upper and lower surfaces.
Fruticose
FruticoseShrubby or hair-like three-dimensional growth form.
Gelatinous
GelatinousJelly-like growth form that swells when wet; characteristic of cyanobacterial lichens.
Gyrophoric acid
A lichen compound that gives a C+ red or pink reaction in the spot test.
Holdfast
Simple attachment structure used by fruticose lichens to anchor to the substrate.
Hymenium
Spore-producing layer inside apothecia.
Isidia
IsidiaCorticated, finger-like outgrowths for asexual reproduction. They have their own cortex, unlike soredia. Singular: isidium.
Isidiate
Bearing isidia.
K test
Chemical spot test using 10% potassium hydroxide (KOH) solution; the most commonly used lichen spot test.
Laminal
On the surface of the thallus, as opposed to marginal (on the edge).
Lecanorine
Apothecia with a rim that contains algae and matches the thallus color.
Lecideine
Apothecia with a dark rim lacking algae.
Leprose
Powdery growth form lacking a cortex entirely.
Lichenometry
Dating rock surfaces by measuring lichen size and applying known growth rates.
Lignicolous
Growing on wood.
Lirellae
LirellaeElongated, line-like fruiting bodies found in genera such as Graphis.
Lobules
Small, lobe-like outgrowths used in asexual reproduction.
Macrolichen
Lichen large enough to see detail with the naked eye, typically foliose or fruticose.
Marginal
On the edge or margin of the thallus or lobes.
Medulla
Loosely woven inner layer of fungal hyphae; the cottony center of the lichen body.
Mycobiont
The fungal partner in a lichen symbiosis.
Nitrophile
Organism that thrives in nitrogen-rich conditions.
Nostoc
A genus of cyanobacteria commonly found as the photobiont in gelatinous lichens.
Oligotroph
Organism adapted to nutrient-poor conditions.
Ostiole
Small opening in a perithecium through which spores escape.
P test
Chemical spot test using paraphenylenediamine; detects fumarprotocetraric and protocetraric acids.
Papillae
Small bumps on the thallus surface.
Perithecia
Flask-shaped, enclosed fruiting bodies embedded in the thallus.
Photobiont
The photosynthetic partner (alga or cyanobacterium) in a lichen.
Podetium
PodetiumStalked, vertical reproductive structure characteristic of Cladonia. Plural: podetia.
Pruinose
Having a frosty or powdery surface coating.
Pseudocyphellae
PseudocyphellaePores in the cortex allowing gas exchange; appear as white spots or lines on the surface.
Rhizines
RhizinesRoot-like bundles of hyphae for attachment, characteristic of foliose lichens.
Rhizomorphs
Cord-like structures made of compacted hyphae.
Saxicolous
Growing on rock.
Soralia
Areas where soredia are produced, appearing as breaks in the cortex. Singular: soralium.
Soredia
SorediaPowdery asexual propagules containing algal cells wrapped in fungal hyphae, lacking a cortex. Singular: soredium.
Sorediate
Bearing soredia.
Squamulose
SquamuloseGrowth form with small, overlapping, scale-like lobes.
Substrate
The surface on which a lichen grows (bark, rock, soil, wood, etc.).
Terricolous
Growing on soil.
Thallus
The entire body of a lichen. Plural: thalli.
TLC
Thin-Layer Chromatography: the gold-standard lab technique for identifying lichen secondary compounds.
Tomentose
Covered with tomentum (a fuzzy, felt-like covering).
Tomentum
Fuzzy, felt-like covering of fine hairs on the lichen surface.
Trebouxia
The most common genus of green algal photobiont in lichens worldwide.
Umbilicate
UmbilicateAttached to the substrate by a single central point, like an umbilical cord (e.g., rock tripe).
Usnic acid
Antibiotic compound produced by Usnea and some other lichen genera.
Veins
Raised ridges on the underside of some lichens, notably Peltigera.
← Back to Identification Guide